Tigress dies in Madhya Pradesh after radio-collar infection. T4, the tigress raised in captivity and known for being part of a successful path-breaking breeding experiment of a translocated tigress, died of an infection caused by its radio-collar at Panna Tiger Reserve, officials said on Friday. The hand-reared tigress was found dead by the forest officials after they detected mortality signal — signal emitted by the radio-collar when animals are inactive for a long period — from T4 at 6.45pm on Thursday, said officials at the Panna Tiger Reserve (PTR).